Simulator · Dog / Cat / Bird / Exotic
How animals see: dog, cat and bird vision camera
Dogs and cats have two types of colour-sensing cone cells, so they see blues and yellows but mix up reds and greens. Birds, many lizards, turtles and goldfish have four, including one tuned to ultraviolet, a colour no human can see. Choose an animal and point your camera: the picture is rebuilt from the measured sensitivity of that animal’s cones.
Loading…
Two cone types: blues and yellows are clear, reds and greens look alike. A red ball, a blue ball and a tennis ball on grass.
How to use it
- Pick an animal. Choose a group (pets, birds, farm, wild, reptiles and fish), then an animal. The camera view changes at once. “You” is always there to compare with.
- Tap a sample at the bottom to see a ready-made scene through the same eyes. The Flower, Budgie and Fruit samples carry real ultraviolet patterns, which a phone camera can’t record. The Night sample shows dim light and body heat.
- Compare (the ‹|› button) splits the picture: your view on the left, the animal’s on the right. Drag the line.
- Full screen turns the page into a camera app; Save downloads the picture.
How each animal sees
Cone peaks are the wavelengths each cone type responds to best, measured in each species’ own retina. Sharpness is converted to the clinic eye-chart scale, where 20/20 is normal human vision.
| Animal | Cone types (peak, nm) | Sharpness | Field of view | Special |
|---|---|---|---|---|
| 🧑 You | 3: 419, 531, 558 | 30 c/° (20/20) | 200° | — |
| 🐶 Dog | 2: 428, 554 | 8 c/° (20/75) | 240° | Red–green colour blind |
| 🐱 Cat | 2: 450, 553 | 8 c/° (20/75) | — | About 5.5× more sensitive in dim light |
| 🐰 Rabbit | 2: 421, 509 | 2 c/° (20/300) | almost 360° | Almost 360° view |
| 🐹 Guinea pig | 2: 429, 529 | 1.65 c/° (20/360) | — | Dichromat |
| 🐹 Hamster | 1: 506 | 0.5 c/° (20/1200) | — | One cone type: no colour |
| 🐀 Rat | 2: 359, 509 | 1 c/° (20/600) | — | Ultraviolet cone |
| 🦜 Budgie | 4: 371, 444, 508, 564 | 10 c/° (20/60) | — | Ultraviolet cone (371 nm) |
| 🦚 Peacock | 4: 424, 458, 505, 567 | 20.6 c/° (20/29) | — | Violet cone; sharp side vision |
| 🐦⬛ Crow | 4: 409, 453, 507, 568 | not measured | — | Violet type (gene evidence) |
| 🐦 Myna | 4: 362, 449, 504, 563 | not measured | — | UV type (gene evidence) |
| 🐦 Sparrow | 4: 362, 449, 504, 563 | 4.88 c/° (20/120) | about 335° | UV type (gene evidence) |
| 🐦 Pigeon | 4: 409, 453, 507, 568 | 12.6 c/° (20/50) | 340° | Violet cone; near 340° view |
| 🐔 Hen | 4: 418, 455, 507, 569 | 6.5 c/° (20/90) | — | Violet cone; flicker to ~87 Hz |
| 🐄 Cow | 2: 435, 553 | 2 c/° (20/300) | — | Dichromat; very low sharpness |
| 🐐 Goat | 2: 444, 552 | not measured | — | Dichromat |
| 🐷 Pig | 2: 439, 556 | not measured | — | Dichromat; UV-clear lens |
| 🐴 Horse | 2: 428, 539 | 23.3 c/° (20/26) | about 330–350° | Sharpest farm animal; blind spot in front |
| 🐯 Tiger | 2: 450, 553 | 8 c/° (20/75) | — | Cat values (not measured) |
| 🦁 Lion | 2: 450, 553 | 8 c/° (20/75) | — | Cat values (not measured) |
| 🐘 Elephant | 2: 419, 552 | 3 c/° (20/200) | — | Same pigments as a colour-blind human |
| 🐒 Monkey | 3: 431, 534, 566 | 26.1 c/° (20/23) | — | Trichromat like humans |
| 🦌 Deer | 2: 455, 537 | 5 c/° (20/120) | — | Dichromat |
| 🐿️ Squirrel | 2: 444, 543 | not measured | — | Yellow lens: no UV |
| 🐠 Goldfish | 4: 356, 447, 537, 623 | 2 c/° (20/300) | — | UV and deep-red cones |
| 🐢 Turtle | 4: 372, 458, 515, 617 | not measured | — | UV cone; red oil droplets |
| 🦎 Bearded dragon | 4: 365, 435, 507, 569 | not measured | — | Parietal eye; UV cone (predicted) |
| 🦎 Gecko | 3: 364, 467, 521 | not measured | — | Colour vision in moonlight |
| 🐍 Ball python | 2: 361, 550 | not measured | — | Infrared heat pits |
c/° = cycles per degree, the finest stripes the eye can resolve. Values are from the sources listed below; where studies disagree, the range is given in the animal’s info panel (ⓘ).
A worked example: why a red ball vanishes on grass
Take the Toys sample: a red ball lying on green grass. Here is how strongly each cone type responds, with white paper counting as 100.
| Cone | Red ball | Grass |
|---|---|---|
| Your blue (S) | 5 | 4 |
| Your green (M) | 5 | 26 |
| Your red (L) | 15 | 26 |
| Dog’s blue (S) | 6 | 3 |
| Dog’s yellow-green (L) | 11 | 24 |
Computed by the same model the camera uses, from the colours in the sample picture.
You see the ball at once because of your red and green cones. The ball gives your red cone 3.0 times the signal of your green cone; the grass gives them about the same (26 and 26). That difference is a strong red–green signal.
A dog has only one cone in that part of the spectrum, so the red–green signal does not exist for it. What is left is brightness and a weaker blue–yellow signal: to a dog the ball is about 46% as bright as the grass and a little less yellow, a dull olive-grey on a yellow-olive lawn. The blue ball is different: it gives the dog’s blue cone 66, against 3 for the grass, while its other cone reads 22 against 24. It shows up as clear blue, which is why a blue toy is easier for a dog to spot on grass than a red one.
How the simulation works
- Light. Each pixel is treated as a surface lit by standard daylight (CIE D65). Its three camera values are turned into the smoothest reflectance curve from 380 to 730 nm that gives exactly those values (Burns’ least-slope method).
- Cones. Each animal’s cones are modelled with the Govardovskii (2000) pigment template at the peak wavelengths measured for that species, filtered by its measured lens and, in birds and lizards, by the coloured oil droplets in its cones. Multiplying light × reflectance × cone sensitivity, wavelength by wavelength, gives each cone’s signal.
- Two cone types (dogs, cats, horses, cattle, rabbits, rodents, snakes): the picture is redrawn using only blues, yellows and greys, choosing for every pixel the colour that gives the animal’s two cones exactly the same signals as the real scene. The animal could not tell the two pictures apart; you see what it can and can’t distinguish.
- One cone type (the Syrian hamster): only brightness is left, so the picture is grey.
- Four cone types (birds, goldfish, turtles, lizards): their three visible cones cover roughly the range yours do, and a camera records only three channels, so visible colours are kept. What they add is ultraviolet. In the samples, where a surface truly reflects UV, magenta is laid over it, the convention used by Vasas and colleagues (2024) for bird-view video. It is false colour, because no screen can emit UV.
- A check against real animals. From the cone data alone, the model predicts the colour that looks grey to a two-cone animal. For dogs it gives 482 nm; trained dogs confused about 480 nm with grey (Neitz, Geist and Jacobs 1989). For guinea pigs it gives 475 nm; guinea pigs confused about 480 nm with white (Jacobs and Deegan 1994). Nothing in the model was tuned to get those matches.
- Sharpness. The picture is blurred by the animal’s measured visual acuity, using the AcuityView method (Caves & Johnsen 2018) and the angle the scene covers. On a phone screen a dog’s view looks only slightly soft, which is accurate; the Eye test sample makes the difference obvious.
What no simulation can show
- What a colour feels like. Nobody can know what yellow is like to a dog. The simulation shows which colours an animal can tell apart, not its experience.
- Ultraviolet through your camera. Phone cameras have a filter that blocks UV, so the live view can’t show it. That is why the samples carry their own UV layer.
- Heat. A snake’s pit organs sense infrared heat, a separate sense from its eyes. Cameras can’t record it, so heat appears only in the Night sample.
- The whole field of view. A horse or rabbit sees almost all the way round. A screen shows one window; the info panel (ⓘ) draws the full field.
- Motion and flicker. Dogs and birds can see faster flicker than you. Your screen can’t show what you can’t see.
- The brain. Animals learn and attend differently. A dog does not mistake its red ball for grass once it smells it.
Common questions
- Do dogs see in black and white?
No. Dogs have two cone types, peaking near 428 and 554 nm, which gives colour vision like a person with red–green colour blindness. They see blues and yellows well; reds, oranges and greens look like shades of yellowish-brown, and blue-green looks grey.
- Can birds see colours we can’t?
Yes. Birds have four cone types, and in parrots such as budgerigars one of them peaks in the ultraviolet, near 371 nm. Budgie feathers carry ultraviolet patterns that play a part in choosing a mate. The Flower and Budgie samples show those patterns in false colour.
- Can cats see in the dark?
Not in total darkness, but in dim light about 5.5 times better than people, mainly thanks to their large pupils and eye optics, plus a reflective layer behind the retina (the tapetum lucidum, which makes their eyes shine). In dim light their vision is grey, like ours.
- Can snakes see heat?
Pythons, boas and pit vipers sense infrared heat with pit organs in their lips or face, not with their eyes. The brain combines the two, so a warm mouse stands out even in the dark.
- Why can’t my phone show ultraviolet?
Camera sensors are built to match human eyes and have a filter that blocks UV and infrared. Your screen can only make red, green and blue light. So any picture of UV, here or anywhere, is false colour.
- Does the camera picture leave my phone?
No. Everything runs in your browser. Nothing is uploaded or saved unless you press Save.
Sources
- Govardovskii VI, Fyhrquist N, Reuter T, Kuzmin DG, Donner K (2000). In search of the visual pigment template. Visual Neuroscience 17:509-528
- Hart NS, Vorobyev M (2005). Modelling oil droplet absorption spectra and spectral sensitivities of bird cone photoreceptors. J Comp Physiol A 191:381-392
- Brettel H, Viénot F, Mollon JD (1997). Computerized simulation of color appearance for dichromats. J Opt Soc Am A 14:2647-2655
- Caves EM, Johnsen S (2018). AcuityView: an R package for portraying the effects of visual acuity on scenes observed by an animal. Methods Ecol Evol 9:793-797
- Burns SA. Generating reflectance curves from sRGB triplets (least slope squared method)
- CIE 015:2018 Colorimetry, 4th ed. (D65 illuminant and 1931 colour-matching functions)
- Vasas V, Lowell MC, Villa J, et al. (2024). Recording animal-view videos of the natural world using a novel camera system and software package. PLoS Biol 22:e3002444 (false-colour conventions)
- Koski MH, Ashman TL (2014). Dissecting pollinator responses to a ubiquitous ultraviolet floral pattern in the wild. Funct Ecol 28:868-877
- Pearn SM, Bennett ATD, Cuthill IC (2001). Ultraviolet vision, fluorescence and mate choice in a parrot, the budgerigar. Proc R Soc B 268:2273-2279
- Pearn SM, Bennett ATD, Cuthill IC (2003). The role of ultraviolet-A reflectance and ultraviolet-A induced fluorescence in the appearance of budgerigar plumage: insights from spectrofluorometry and reflectance spectrophotometry. Proc R Soc B 270:859-865
- Siitari H, Honkavaara J, Viitala J (1999). Ultraviolet reflection of berries attracts foraging birds. Proc R Soc B 266:2125-2129
- Middleton R, Tunstad SA, Knapp A, et al. (2024). Self-assembled, disordered structural color from fruit wax bloom. Sci Adv 10:eadk4219
- Neitz J, Geist T, Jacobs GH (1989). Color vision in the dog. Vis Neurosci 3:119-125
- Jacobs GH, Deegan JF, Crognale MA, Fenwick JA (1993). Photopigments of dogs and foxes and their implications for canid vision. Vis Neurosci 10:173-180
- McDowell RJ, Didikoglu A, Woelders T, et al. (2024). Beyond Lux: methods for species and photoreceptor-specific quantification of ambient light for mammals. BMC Biol 22:257 (pigment peaks and lens spectra)
- Douglas RH, Jeffery G (2014). The spectral transmission of ocular media suggests ultraviolet sensitivity is widespread among mammals. Proc R Soc B 281:20132995
- Murphy CJ, Mutti DO, Zadnik K, Ver Hoeve J (1997). Effect of optical defocus on visual acuity in dogs. Am J Vet Res 58:414-418
- Miller PE, Murphy CJ (1995). Vision in dogs. J Am Vet Med Assoc 207:1623-1634
- Lind O, Milton I, Andersson E, Jensen P, Roth LSV (2017). High visual acuity revealed in dogs. PLoS One 12:e0188557
- Coile DC, Pollitz CH, Smith JC (1989). Behavioral determination of critical flicker fusion in dogs. Physiol Behav 45:1087-1092
- Healy K, McNally L, Ruxton GD, Cooper N, Jackson AL (2013). Metabolic rate and body size are linked with perception of temporal information. Anim Behav 86:685-696
- Guenther E, Zrenner E (1993). The spectral sensitivity of dark- and light-adapted cat retinal ganglion cells. J Neurosci 13:1543-1550
- Jacobs GH, Neitz J (1986). Spectral sensitivity of cat cones to rapid flicker. Exp Brain Res 62:446-448
- Kang I, Reem RE, Kaczmarowski AL, Malpeli JG (2009). Contrast sensitivity of cats and humans in scotopic and mesopic conditions. J Neurophysiol 102:831-840
- Duffy KR, Bear MF, Patel NB, Das VE, et al. (2023). Human deprivation amblyopia: treatment insights from animal models. Front Neurosci 17:1249466 (cat acuity, citing Giffin & Mitchell 1978)
- Ollivier FJ, Samuelson DA, Brooks DE, Lewis PA, Kallberg ME, Komáromy AM (2004). Comparative morphology of the tapetum lucidum (among selected species). Vet Ophthalmol 7:11-22
- Nuboer JFW, van Nuys WM, Wortel JF (1983). Cone systems in the rabbit retina revealed by ERG-null-detection. J Comp Physiol 151:347-352
- Peiffer RL Jr, Pohm-Thorsen L, Corcoran K (1994). Models in ophthalmology and vision research. In: The Biology of the Laboratory Rabbit, 2nd ed., pp. 409-433
- Jacobs GH, Deegan JF (1994). Spectral sensitivity, photopigments, and color vision in the guinea pig (Cavia porcellus). Behav Neurosci 108:993-1004
- Jnawali A, Puri S, Frishman LJ, Ostrin LA (2021). Visual function in guinea pigs: behavior and electrophysiology. Clin Exp Optom 104:523-531
- Calderone JB, Jacobs GH (1999). Cone receptor variations and their functional consequences in two species of hamster. Vis Neurosci 16:53-63
- Emerson VF, Chalupa LM, Thompson ID, Talbot RJ (1982). Behavioural, physiological, and anatomical consequences of monocular deprivation in the golden hamster. Exp Brain Res 45
- Jacobs GH, Neitz J, Deegan JF (1991). Retinal receptors in rodents maximally sensitive to ultraviolet light. Nature 353:655-656
- Prusky GT, West PWR, Douglas RM (2000). Behavioral assessment of visual acuity in mice and rats. Vision Res 40:2201-2209
- Prusky GT, Harker KT, Douglas RM, Whishaw IQ (2002). Variation in visual acuity within pigmented, and between pigmented and albino rat strains. Behav Brain Res 136:339-348
- Bowmaker JK, Heath LA, Wilkie SE, Hunt DM (1997). Visual pigments and oil droplets from six classes of photoreceptor in the retinas of birds. Vision Res 37:2183-2194
- Lind O, Mitkus M, Olsson P, Kelber A (2014). Ultraviolet vision in birds: the importance of transparent eye media. Proc R Soc B 281:20132209
- Lind O, Kelber A (2011). The spatial tuning of achromatic and chromatic vision in budgerigars. J Vis 11(7):2
- Boström JE, Haller NK, Dimitrova M, Ödeen A, Kelber A (2017). The flicker fusion frequency of budgerigars (Melopsittacus undulatus) revisited. J Comp Physiol A 203:15-22
- Vorobyev M (2003). Coloured oil droplets enhance colour discrimination. Proc R Soc B 270:1255-1261
- Vasas V, Lowell MC, Villa J, et al. (2024). Recording animal-view videos of the natural world using a novel camera system and software package. PLoS Biol 22:e3002444
- Hart NS (2002). Vision in the peafowl (Aves: Pavo cristatus). J Exp Biol 205:3925-3935
- Ödeen A, Håstad O (2013). The phylogenetic distribution of ultraviolet sensitivity in birds. BMC Evol Biol 13:36
- Kane SA, Wang Y, Fang R, Lu Y, Dakin R (2019). How conspicuous are peacock eyespots and other colorful feathers in the eyes of mammalian predators? PLoS One 14:e0210924
- Loyau A, Gomez D, Moureau B, Théry M, Hart NS, Saint Jalme M (2007). Iridescent structurally based coloration of eyespots correlates with mating success in the peacock. Behav Ecol 18:1123-1131
- Dakin R, Montgomerie R (2013). Eye for an eyespot: how iridescent plumage ocelli influence peacock mating success. Behav Ecol 24:1048-1057
- Ödeen A, Håstad O (2003). Complex distribution of avian color vision systems revealed by sequencing the SWS1 opsin from total DNA. Mol Biol Evol 20:855-861
- Ödeen A, Håstad O, Alström P (2011). Evolution of ultraviolet vision in the largest avian radiation – the passerines. BMC Evol Biol 11:313
- Fernández-Juricic E, O’Rourke C, Pitlik T (2010). Visual coverage and scanning behavior in two corvid species. J Comp Physiol A 196:879-888
- Hart NS, Partridge JC, Cuthill IC (1998). Visual pigments, oil droplets and cone photoreceptor distribution in the European starling (Sturnus vulgaris). J Exp Biol 201:1433-1446
- Dolan T, Fernández-Juricic E (2010). Retinal ganglion cell topography of five species of ground-foraging birds. Brain Behav Evol 75:111-121
- Fernández-Juricic E, Gall MD, Dolan T, Tisdale V, Martin GR (2008). The visual fields of two ground-foraging birds, House Finches and House Sparrows, allow for simultaneous foraging and anti-predator vigilance. Ibis 150:779-787
- Hahmann U, Güntürkün O (1993). The visual acuity for the lateral visual field of the pigeon. Vision Res 33:1659-1664
- Martin GR, Young SR (1983). The retinal binocular field of the pigeon. Vision Res 23:911-915
- Gover N, Jarvis JR, Abeyesinghe SM, Wathes CM (2009). Stimulus luminance and the spatial acuity of domestic fowl. Vision Res 49:2747-2753
- Lisney TJ, Rubene D, Rózsa J, et al. (2011). Behavioural assessment of flicker fusion frequency in chicken. Vision Res 51:1324-1332
- Jacobs GH, Deegan JF, Neitz J (1998). Photopigment basis for dichromatic color vision in cows, goats, and sheep. Vis Neurosci 15:581-584
- Rehkämper G, Görlach A (1998). Visual identification of small sizes by adult dairy bulls. J Dairy Sci 81:1574-1580
- Neitz J, Jacobs GH (1989). Spectral sensitivity of cones in an ungulate. Vis Neurosci 2:97-100
- Ahnelt PK, Schubert C, Kübber-Heiss A, Schiviz A, Anger E (2006). Independent variation of retinal S and M cone photoreceptor topographies: a survey of four families of mammals. Vis Neurosci 23:429-435
- Carroll J, Murphy CJ, Neitz M, Ver Hoeve JN, Neitz J (2001). Photopigment basis for dichromatic color vision in the horse. J Vis 1:80-87
- Timney B, Keil K (1992). Visual acuity in the horse. Vision Res 32:2289-2293
- Harman AM, Moore S, Hoskins R, Keller P (1999). Horse vision and an explanation for the visual behaviour originally explained by the ‘ramp retina’. Equine Vet J 31:384-390
- Rørvang MV, Nielsen BL, McLean AN (2020). Sensory abilities of horses and their importance for equitation science. Front Vet Sci 7:633
- Gunter R (1951). The absolute threshold for vision in the cat. J Physiol 114:8-15
- Yokoyama S, Takenaka N, Agnew DW, Shoshani J (2005). Elephants and human color-blind deuteranopes have identical sets of visual pigments. Genetics 170:335-344
- Shyan-Norwalt MR, Peterson J, Milankow King B, Staggs TE, Dale RHI (2010). Initial findings on visual acuity thresholds in an African elephant (Loxodonta africana). Zoo Biol 29:30-35
- Pettigrew JD, Bhagwandin A, Haagensen M, Manger PR (2010). Visual acuity and heterogeneities of retinal ganglion cell densities and the tapetum lucidum of the African elephant (Loxodonta africana). Brain Behav Evol 75:251-261
- Hárosi FI (1987). Cynomolgus and rhesus monkey visual pigments. J Gen Physiol 89:717-743
- Ridder WH, et al. (2019). Comparison of contrast sensitivity in macaque monkeys and humans. Vis Neurosci 36:E008
- Jacobs GH, Deegan JF, Neitz J, Murphy BP, Miller KV, Marchinton RL (1994). Electrophysiological measurements of spectral mechanisms in the retinas of two cervids: white-tailed deer and fallow deer. J Comp Physiol A 174:551-557
- (2022). Estimation of visual discrimination in the white-tailed deer by behavioral assay. Am Midl Nat 187:90
- Hogg C, et al. (2011). Arctic reindeer extend their visual range into the ultraviolet. J Exp Biol 214:2014-2019
- Blakeslee B, Jacobs GH, Neitz J (1988). Spectral mechanisms in the tree squirrel retina. J Comp Physiol A 162:773-780
- Palacios AG, Varela FJ, Srivastava R, Goldsmith TH (1998). Spectral sensitivity of cones in the goldfish. Vision Res 38:2135-2146
- Bowmaker JK, Thorpe A, Douglas RH (1991). Ultraviolet-sensitive cones in the goldfish. Vision Res 31:349-352
- Neumeyer C (1992). Tetrachromatic color vision in goldfish. J Comp Physiol A 171:639-649
- Neumeyer C (2003). Wavelength dependence of visual acuity in goldfish. J Comp Physiol A 189:811-821
- Loew ER, Govardovskii VI (2001). Photoreceptors and visual pigments in the red-eared turtle, Trachemys scripta elegans. Vis Neurosci 18:753-757
- Yewers MS, McLean CA, Moussalli A, et al. (2015). Spectral sensitivity of cone photoreceptors and opsin expression in two colour-divergent lineages of the lizard Ctenophorus decresii. J Exp Biol 218:1556-1563
- Barbour HR, Archer MA, Hart NS, et al. (2002). Retinal characteristics of the ornate dragon lizard, Ctenophorus ornatus. J Comp Neurol 450:334-344
- Solessio E, Engbretson GA (1993). Antagonistic chromatic mechanisms in photoreceptors of the parietal eye of lizards. Nature 364:442-445
- Su CY, Luo DG, Terakita A, et al. (2006). Parietal-eye phototransduction components and their potential evolutionary implications. Science 311:1617-1621
- Foà A, Basaglia F, Beltrami G, et al. (2009). Orientation of lizards in a Morris water-maze: roles of the sun compass and the parietal eye. J Exp Biol 212:2918-2924
- Kojima D, Okano T, Fukada Y, et al. (1992). Cone visual pigments are present in gecko rod cells. PNAS 89:6841-6845
- Loew ER (1994). A third, ultraviolet-sensitive, visual pigment in the Tokay gecko (Gekko gekko). Vision Res 34:1427-1431
- Roth LSV, Kelber A (2004). Nocturnal colour vision in geckos. Proc R Soc B 271(Suppl 6):S485-S487
- Roth LSV, Lundström L, Kelber A, Kröger RHH, Unsbo P (2009). The pupils and optical systems of gecko eyes. J Vis 9(3):27
- Abramjan A, Baranová V, Frýdlová P, Landová E, Frynta D (2020). Ultraviolet reflectance and pattern properties in leopard geckos. Behav Processes 173:104060
- Davies WL, Cowing JA, Bowmaker JK, et al. (2009). Shedding light on serpent sight: the visual pigments of henophidian snakes. J Neurosci 29:7519-7525
- Sillman AJ, Carver JK, Loew ER (1999). The photoreceptors and visual pigments in the retina of a boid snake, the ball python (Python regius). J Exp Biol 202:1931-1938
- Gracheva EO, Ingolia NT, Kelly YM, et al. (2010). Molecular basis of infrared detection by snakes. Nature 464:1006-1011
- Hartline PH, Kass L, Loop MS (1978). Merging of modalities in the optic tectum: infrared and visual integration in rattlesnakes. Science 199:1225-1229
- Bakken GS, Krochmal AR (2007). The imaging properties and sensitivity of the facial pits of pitvipers as determined by optical and heat-transfer analysis. J Exp Biol 210:2801-2810